Default header image

Grad Students

There have been so many students pass through our doors over the years, both undergraduates and graduates, including students supervised by both Dr David Pengelly and Dr. Steve Marshall. Here are Dr. Marshall’s current graduate students (alphabetical). For a list of Dr Marshall’s previous grad students, check out his page here.

Gustavo Borges Ferro, PhD Candidate

Gustavo came to us in 2015 after having worked on the Micropezidae in Brazil, and is getting very close to finishing up his PhD here. His research focuses on the generic relationships within the subfamily Taeniapterinae, and has completed several revisions of micropezid genera along the way.


To see a list of Gustavo’s publications, please check out his maintained cv in English and Português.


Gregory Kuwahara, MSc Student

Gregory photographing insects.

Gregory started working in the Insect Collection in 2018, helping Dr. Paiero curate the diving beetle section of the Collection. He then went on to do an undergraduate project with Dr. Marshall during his undergraduate degree in Zoology on a neotropical genus of Sphaerocerid flies (Chespiritos) and is now working on a revision of the Australian flies in the genus Howickia. Contact Gregory at


Kate Lindsay, MSc Student

Kate checking out New Zealand

Kate began working in the collection in 2017, when she started a work study during her undergrad in Zoology. She reviewed the Laphria (Diptera: Asilidae) of Eastern Canada for her undergrad project and shortly after undertook a project reviewing a small species complex from the genus Curtonotum (Diptera: Curtonotidae). She is currently doing her Masters on the micropezid genus Scipopus (Diptera: Micropezidae).


  • Lindsay, K.G. and Marshall, S.A. 2019. Laphria (Diptera: Asilidae) of Ontario, with a key to the eastern Canadian species of Laphriini and Dasylechia. Canadian Journal of Arthropod Identification 37: 91pp. doi:10.3752/cjai.2019.37
  • Lindsay, K., Klymko, J. and Marshall, S. A. 2019. Revision of the Central American and Mexican species of the Curtonotum murinum species complex (Diptera: Curtonotidae). Zootaxa (In Press)


Nichelle Lomas, PhD Candidate

Nichelle strumming the banjo!

Nichelle joined the lab September 2009 after graduating with an undergrad zoology degree from U of G. She is a student of both Dr. Marshall and Dr. Youbin Zheng undertaking a OMAFRA funded project unusual for the systematics lab as her flies are (hopefully) still living. She worked on an experimental project studying the use of saprophagous fly larvae to convert chicken manure into fertilizer, breeding colonies of several fly species to test their effectiveness as waste converters. Nichelle’s research is the entomological side of a larger project by Ivan Milin, president of ecospace engineering ( Contact her at


Tiffany Yau, PhD Student

Tiffany photographing a Flatidae in Costa Rica.

Tiffany started as a work study student in her 3rd year undergrad and did an undergraduate project with Steve in 2013, utilizing her artistic skills to examine several undescribed genera of Sphaeroceridae, focusing on the morphology of the genitalia. She finished her M.Sc. here revising a genus of Sphaeroceridae (Bromeloecia) and is continuing her research on the taxonomy and systematics of the the Pseudopomyzidae (Diptera) in her Ph.D. Contact her at


  • Entomological Society of Ontario
  • Entomological Society of Canada


  • Marshall, S. A., & Yau, T. (2014). Paramosina, a new genus of high Andean Limosininae (Diptera: Sphaeroceridae). Zootaxa, 3872(4), 393-397.
  • Yau T. and Marshall S.A. (2015). Enzymatic clearing agents as an alternative approach to macerating Diptera specimens. Fly Times (55): 11–12
  • Yau, T., & Marshall, S. A. (2018). A revision of the genus Bromeloecia Spuler (Diptera: Sphaeroceridae: Limosininae). Zootaxa, 4445(1), 1-115.
  • Borkent, A., Brown, B. V., Adler, P. H., Amorim, D. D. S., Barber, K., Bickel, D., … & Capellari, R. S. (2018). Remarkable fly (Diptera) diversity in a patch of Costa Rican cloud forest: Why inventory is a vital science. Zootaxa, 4402(1), 53-90.
  • Brown, B. V., Borkent, A., Adler, P. H., de Souza Amorim, D., Barber, K., Bickel, D., … & Capellari, R. S. (2018). Comprehensive inventory of true flies (Diptera) at a tropical site. Communications biology, 1(1), 21.
  • Marshall, S.A. and Yau, T. (2019) Coprosylia bisetosa, a new genus and species of limosinine sphaerocerid (Diptera: Sphaeroceridae) phoretic on dung beetles in Borneo. Raffles Bulletin of Zoology, 67: 183-188.

Other Projects

  • Yau T. and Marshall, S.A. A revision of the subgenus Pseudopomyza (Rhinopomyzella) (Diptera, Pseudopomyzidae) (Pending)
  • Marshall, S.A. and Yau, T. Stipulosina, a new genus of Limosininae (Diptera: Sphaeroceridae) associated with bamboo stipules in Ecuador. (Pending)


Nur Athiqah Md Yosuf, PhD Candidate

Nur visiting the United States National Museum

Nur came to work here with us in 2016 and is currently finishing up her thesis revising the Eurybatinae (Diptera: Micropezidae) and it’s relationships with other micropezid subfamilies.