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Grad Students

There have been so many students pass through our doors over the years, both undergraduates and graduates, including students supervised by both Dr David Pengelly and Dr. Steve Marshall. Following his retirement, Dr. Marshall’s graduate students have completed their research, although several graduate students within the department utilize the collection as a reference collection and research lab. For a list of Dr Marshall’s previous grad students, check out his page here. A few of our recent graduates are listed below!


Gregory Kuwahara, MSc Graduate

Gregory photographing insects.

Gregory started working in the Insect Collection in 2018, helping Dr. Paiero curate the diving beetle section of the Collection. He then went on to do an undergraduate project with Dr. Marshall during his undergraduate degree in Zoology on a neotropical genus of Sphaerocerid flies (Chespiritos), completed his revision of Howickia (Sphaeroceridae: Limosininae), and is now heavily involved with Dr Marshall’s research in several other genera of Sphaeroceridae. Contact Gregory at


  • Kuwahara, Gregory K. & Stephen A. Marshall. 2020. A revision of the Neotropical genus Chespiritos (Diptera: Sphaeroceridae: Limosininae). European Journal of Entomology 117: 164-189. 10.14411/eje.2020.019
  • Kuwahara, Gregory K. & Stephen A. Marshall. 2022. A revision of the Australian species of Howickia Richards (Diptera: Sphaeroceridae: Limosininae). Zootaxa 5192(1): 1-152. 10.11646/zootaxa.5192.1.1


Kate Lindsay, MSc Graduate

Kate checking out New Zealand

Kate began working in the collection in 2017, when she started a work study during her undergrad in Zoology. She reviewed the Laphria (Diptera: Asilidae) of Eastern Canada for her undergrad project and shortly after undertook a project reviewing a small species complex from the genus Curtonotum (Diptera: Curtonotidae). She finished up her Masters in 2021 on the micropezid genus Scipopus (Diptera: Micropezidae), and is currently involved a revision of a related genus.


  • Lindsay, K.G. and Marshall, S.A. 2023. A revision of Scipopus Enderlein including the subgenera Scipopus s. str., Phaeopterina Frey and Parascipopus subgen. nov. (Diptera, Micropezidae, Taeniapterinae). European Journal of Taxonomy 904: 1-189. 10.5852/ejt.2023.904.2323
  • Lindsay, K.G. and Marshall, S.A. 2019. Laphria (Diptera: Asilidae) of Ontario, with a key to the eastern Canadian species of Laphriini and Dasylechia. Canadian Journal of Arthropod Identification 37: 91pp. doi:10.3752/cjai.2019.37
  • Lindsay, K., Klymko, J. and Marshall, S. A. 2019. Revision of the Central American and Mexican species of the Curtonotum murinum species complex (Diptera: Curtonotidae). Zootaxa (In Press)


Tiffany Yau, PhD Graduate

Tiffany photographing a Flatidae in Costa Rica.

Tiffany started as a work study student in her 3rd year undergrad and did an undergraduate project with Steve in 2013, utilizing her artistic skills to examine several undescribed genera of Sphaeroceridae, focusing on the morphology of the genitalia. She finished her PhD on the taxonomy and systematics of the the Pseudopomyzidae (Diptera) in 2023 and continues to wrap up various projects she was involved in during her graduate studies. Contact her at


  • Entomological Society of Ontario
  • Entomological Society of Canada


  • Marshall, S. A., & Yau, T. (2014). Paramosina, a new genus of high Andean Limosininae (Diptera: Sphaeroceridae). Zootaxa, 3872(4), 393-397.
  • Yau T. and Marshall S.A. (2015). Enzymatic clearing agents as an alternative approach to macerating Diptera specimens. Fly Times (55): 11–12
  • Yau, T., & Marshall, S. A. (2018). A revision of the genus Bromeloecia Spuler (Diptera: Sphaeroceridae: Limosininae). Zootaxa, 4445(1), 1-115.
  • Borkent, A., Brown, B. V., Adler, P. H., Amorim, D. D. S., Barber, K., Bickel, D., … & Capellari, R. S. (2018). Remarkable fly (Diptera) diversity in a patch of Costa Rican cloud forest: Why inventory is a vital science. Zootaxa, 4402(1), 53-90.
  • Brown, B. V., Borkent, A., Adler, P. H., de Souza Amorim, D., Barber, K., Bickel, D., … & Capellari, R. S. (2018). Comprehensive inventory of true flies (Diptera) at a tropical site. Communications biology, 1(1), 21.
  • Marshall, S.A. and Yau, T. (2019) Coprosylia bisetosa, a new genus and species of limosinine sphaerocerid (Diptera: Sphaeroceridae) phoretic on dung beetles in Borneo. Raffles Bulletin of Zoology, 67: 183-188.

Other Projects

  • Yau T. and Marshall, S.A. A revision of the subgenus Pseudopomyza (Rhinopomyzella) (Diptera, Pseudopomyzidae) (Pending)
  • Marshall, S.A. and Yau, T. Stipulosina, a new genus of Limosininae (Diptera: Sphaeroceridae) associated with bamboo stipules in Ecuador. (Pending)